Russia has lost 1,110 fighters and 66 artillery systems in Ukraine over the past day, Kyiv's armed forces said on Monday, after Moscow said its troops had taken two more villages in the east of the war-torn country.

Russia has sustained a total of 543,810 casualties—referring to soldiers both killed and injured—in Ukraine since February 2022, Ukraine's military said in an update posted to social media. Moscow has lost just under 14,600 artillery systems since the Kremlin launched its full-scale invasion more than two years ago, according to Kyiv's statistics.

Ukraine's General Staff also said on Monday that Russia had lost 115 vehicles of varying types since early on Sunday, including 19 tanks and 26 armored personnel vehicles.

Newsweek could not independently verify these numbers and has reached out to the Russian Defense Ministry for comment via email.

Casualty counts and battlefield losses are murky during active conflicts, and experts urge caution when dealing with tallies offered up by either party in a war.

The British government estimated in late May that Russia's total casualties had likely reached 500,000. Throughout May 2024, Moscow's average daily casualty count stood at more than 1,200, the U.K. Defense Ministry said, marking the highest average number of daily losses for Russia of the full-scale war.

Russia was still plugging away with recruitment efforts to replenish the ranks hit by Moscow's "ongoing attritional offensive" across hundreds of miles of front lines, the U.K. government added at the time.

Ukraine has said Russia is able to pull up to 30,000 new soldiers each month into its ranks, propping up the bloody and casualty-heavy tactics Moscow has used against Kyiv's forces to make slow but consistent territorial gains.

While launching a cross-border offensive from Russia's Belgorod region into Ukraine's northeastern Kharkiv region in early May, Moscow has doubled down on its efforts to inch westward in Ukraine's eastern Donetsk region.

Ukraine's military has repeatedly reported the heaviest fighting occurs east of the strategic Donetsk city of Pokrovsk as Russia advances west of the former Ukrainian stronghold of Avdiivka. Moscow has controlled Avdiivka since February.

On Sunday, Russia's Defense Ministry said its forces had claimed two villages in the Donetsk region. Its forces now control Spirne, a settlement in the east of Donetsk and not far from the Luhansk region, Moscow said. It has also captured Novooleksandrivka, a village east of Pokrovsk and northwest of Avdiivka.

Ukraine's military said on Monday that its troops had "repelled" eight Russian assaults east of the Donetsk settlement of Siversk in the past day, including around Spirne.

Ukraine fought off 42 Russian attacks east of Pokrovsk, including assaults around Novooleksandrivka, Kyiv's military said in an operational update.